*=undergraduate student author; ^=graduate student author; **=former student
2023
Siebert, S. J., T. J. Samojedny Jr.*, R. B. Boneschans, H. W. Ostmann*, C. V. Clarke*, and N. Rajakaruna. Searching for new nickel hyperaccumulating plants: Introducing X-Ray Fluorescence (XRF) analysis of herbarium specimens to southern Africa (To be submitted: South African Journal of Botany)
Van Staden, N., Siebert, F., Van Zijl, G., Rajakaruna, N., and S. Siebert. Plant diversity patterns of geologically distinct, semi-arid savanna mountain landscapes of Griqualand West, South Africa (Plant Ecology and Diversity; under revision)
102. Siebert, S. J., S. Palacio, A. L. Luzuriaga, G. Maggs-Kölling, E. Marais, S. Matesanz, M. Prieto, Y. Pueyo, N. Rajakaruna, A. M. Sánchez, and S. Claassens. GYPWORLD Africa: Setting an agenda for gypsum ecosystem research in southern Africa (South African Journal of Science; in press)
101. Picó, F. X., R. J. Abbott, L. D. Llambi, N. Rajakaruna, A. S. T. Papadopulos, and L. Nagy. 2023. Introduction to Special Issue: The ecology and evolution of plants in extreme environments. Plant Ecology and Diversity 15: 179-182; https://doi.org/10.1080/17550874.2022.2164703 (PDF)
100. Samojedny, T. J. Jr.*, C. Garnica-Díaz, D. Grossenbacher, G. C. Adamidis, P. G. Dimitrakopoulos, S. J. Siebert, M. J. Spasojevic, C. M. Hulshof, N. Rajakaruna. 2023. Specific leaf area is lower on ultramafic than on neighbouring non-ultramafic soils. Plant Ecology and Diversity 15: 243-253; https://doi.org/10.1080/17550874.2022.2160673 (PDF)
99. Garnica-Díaz C, Berazaín Iturralde R, Cabrera B, Calderón Morales E, Felipe Tamé FL, García R, Gómez Hechavarría JL, Guimarães AF, Medina E, Paul ALD, Rajakaruna N, Restrepo C, Siebert SJ, van den Berg E, Van der Ent A, Velasquez G, and CM Hulshof. 2023. Global plant ecology of tropical ultramafic ecosystems. The Botanical Review (https://doi.org/10.1007/s12229-022-09278-2; PDF)
2022
98. Eskandari, B. S., M. S. Gahrouei, R. S. Boyd, N. Rajakaruna, R. Ghasemi. 2022. Physiological responses to lead and PEG-simulated drought stress in metallicolous and non-metallicolous Matthiola (Brassicaceae) species from Iran. South African Journal of Botany 150: 1011-1021(PDF)
97. Mulroy, M^., Fryday, A. M., Gersoff, A*., Dart, J., Næsborg, R. R., and N. Rajakaruna. 2022. Lichens of Ultramafic Substrates in North America: A Review. Botany; https://doi.org/10.1139/cjb-2021-0187 (PDF)
96. Samojedny, T.J.*, Devlin, M.**, Shane, R.*, and Rajakaruna, N. 2022. The effects of nitrogen enrichment on low nutrient environments: Insights from studies of serpentine soil-plant relations. In: Naeem, M., Bremont, J.F.J., Ansari, A.A., Gill, S.S. (eds) Agrochemicals in Soil and Environment. Springer, Singapore. https://doi.org/10.1007/978-981-16-9310-6_13 (PDF)
2021
95. Fernando, D. R., A. van der Ent, M., H. A. S. Weerasinghe, D. S. A. Wijesundera, G. W. A. R. Fernando, A. E. Fernando, M. C.M. Iqbal, C. H. Miranda*, J. M. Gosse*, Y. A. S. Samithri and N. Rajakaruna. 2021. Assessment of plant diversity and foliar chemistry on the Sri Lankan ultramafics reveals inconsistencies in the metal hyperaccumulator trait. Ecological Research; https://doi.org/10.1111/1440-1703.12282 (PDF; Supporting Files – Figure S1; Table S1)
Graphical Abstract: A field survey of all known Sri Lankan ultramafic outcrops revealed high levels of plant diversity (132 species in 44 families) and rediscovered the facultative nickel hyperaccumulator Rinorea bengalensis (Violaceae).
94. Adhikari, S., J. M. Silva, N. Rajakaruna, and S. J. Siebert. 2021. Influence of land use and topography on distribution and bioaccumulation of potentially toxic metals in soil and plant leaves: A case study from Sekhukhuneland, South Africa. Science of the Total Environment 806, Part 2,150659; https://doi.org/10.1016/j.scitotenv.2021.150659 (PDF)
Graphical Abstract:
93. Fryday, A. M., Siebert, S. J., N. Rajakaruna. 2021. Scoring a lichen hat-trick. Veld & Flora (in press; June Issue, p. 26-31)
92. Naikoo, M. I., Khan F, A., Noureldeen, A., Rinklebe, J., Sonne, C., Rajakaruna, N., Ahmad, P. 2021. Biotransfer, Bioaccumulation and Detoxification of Nickel along the Soil- Faba Bean-Aphid-Ladybird Food Chain. Science of the Total Environment 785; DOI: https://doi.org/10.1016/j.scitotenv.2021.147226 (PDF)
Graphical Abstract:
91. Yu, A. C., M. Reinhart*, R. Hunter*, K. Lu, K. Maikawa, N. Rajakaruna, J. Acosta, Stubler, C. Appel, E. Appel. 2021. Seasonal impact of phosphate-based fire retardants on soil chemistry following prophylactic treatment of vegetation. Environmental Science and Technology 55: 2316-2323 (PDF)
2020
90. Fryday, A. M., Medeiros, I.**, Pope, N.**, Siebert, S. and N. Rajakaruna. Burrowsia, a new genus of lichenized fungi (Caliciaceae), plus the new species B. cataractarum and Scoliciosporum fabisporum, from Mpumalanga, South Africa. South African Journal of Botany 121: 471-481 (PDF)
89. Ferrero, A.*, Walsh, P.*, and N. Rajakaruna. 2020. The physiology, genetics, adaptive significance and biotechnology of Ni-hyperaccumulating plants. In: Physiological and Biotechnological Aspects of Extremophiles, pp. 327-347. Richa Salwan, Vivek Sharma (Eds.), Elsevier Publishers (invited book chapter; PDF)
2019
88. Gunarathne, V., A. U. Rajapaksha, M. Vithanage, N. Adassooriya, A. Cooray, S. Liyanage, B. Athapattu, N. Rajakaruna, A. D. Igalavithana, D. Hou, D. S. Alessi, and Y. S. Ok. 2019. Heavy Metal Dissolution Mechanisms from Electrical Industrial Sludge. Science of the Total Environment (PDF)
87. Paukov, A., A. Teptina, M. Morozova, E. Kruglova, S. Favero-Longo, C. Bishop* and N. Rajakaruna. The effects of edaphic and climatic factors on secondary lichen chemistry—a case study using saxicolous lichens. Diversity (PDF)
86. Naikoo, M. I., M. I. Dar, F. A. Khan, F. Raghib, and N. Rajakaruna. Trophic transfer and bioaccumulation of lead along soil – plant – aphid – ladybird food chain. Environmental Science and Pollution Research 26: 23460–23470 (PDF)
85. Gunarathne, V., N. Rajakaruna, U. Gunarathne, J. K. Biswas, Z. A. Raposo*, and M. Vithanage. 2019. Influence of soil water content and soil amendments on trace metal release and seedling growth in serpentine soil. Journal of Soils and Sediments (PDF)
84. Ul Kalam, S., F. Naushin, F. A. Khan, and N. Rajakaruna. 2019. Long term phytoremediating abilities of Dalbergia sissoo Roxb. (Fabaceae). SN Applied Sciences 1: 501 (PDF)
2018
83. Echevarria, G., A. J. M. Baker, R. S. Boyd, A. van der Ent, T. Mizuno, N. Rajakaruna, S. Sakaguchi, and A. Bani. 2018. A global forum on ultramafic ecosystems: from ultramafic ecology to rehabilitation of degraded environments. Ecological Research 33: 517-522. (PDF)
82. Kay, K.M., Woodhouse, S.^, Smith, B.A., Pope, N.**, and Rajakaruna, N. Sympatric serpentine endemic Monardella (Lamiaceae) species maintain habitat differences despite hybridization. Molecular Ecology 27: 2302-2316. [PDF; Master’s Thesis research of Suzie Woolhouse (SJSU) and Brett Smith (UCSC)]; COVER IMAGE
81. Teptina, A., A. Paukov, and N. Rajakaruna. 2018. Ultramafic vegetation and soils in the circumboreal region of the Northern Hemisphere. Ecological Research 33: 609-628. (PDF)
80. Favero Longo, S. E., E. Matteucci, P. Giordani, A. Paukov, and N. Rajakaruna. 2018. Diversity and functional traits of lichens in ultramafic areas: a literature-based worldwide analysis integrated by field data at the regional scale. Ecological Research 33: 593-608. (PDF)
79. Ghasemi, R., H. Share, R. Sharifi, R. S. Boyd, N. Rajakaruna. 2018. Inducing Ni sensitivity in the Ni hyperaccumulator plant Alyssum inflatum Nyárády (Brassicaceae) by transforming with CAX1, a vacuolar membrane calcium transporter. Ecological Research 33: 737:747. (PDF)
78. Rajakaruna, N. 2018. Lessons on evolution from the study of edaphic specialization. The Botanical Review 84: 39-78; DOI 10.1007/s12229-017-9193-2 (PDF)
2017
77. Venter, A., S. J. Siebert, N. Rajakaruna, S. Barnard, A. Levanets, A. Ismail, M. Allam, B. Peterson, T. Sanko. 2017. Biological crusts of serpentine and non-serpentine soils from the Barberton Greenstone Belt of South Africa. Ecological Research 33: 629-640. (PDF)
76. Siebert, S. J., Schutte, N. C., Bester, S. P., Komape, D. M., and N. Rajakaruna. 2017. Senecio conrathii N.E.Br. (Asteraceae), a new hyperaccumulator of nickel from serpentinite outcrops of the Barberton Greenstone Belt, South Africa. Ecological Research 33: 651-658. (PDF)
75. Galey, M. L.**, A. van der Ent, M. C.M. Iqbal, and N. Rajakaruna. 2017. Serpentine geoecology of South and Southeast Asia. Botanical Studies 58: 18; DOI 10.1186/s40529-017-0167-9 (PDF; undergraduate Independent Study of Miranda Galey)
74. Seneviratne, M. N. Rajakaruna, M. Rizwan, S. Madawala, Y. S. Ok, and M. Vithanage. 2017. The effects of heavy metal-induced oxidative stress on seed germination and seedling development: A critical review. Environmental Geochemistry and Health; DOI: 10.1007/s10653-017-0005-8 (PDF)
73. Medeiros, I. D.*, Mathieson, A. C., and N. Rajakaruna. 2017. Heavy Metals in Seaweeds from a Polluted Estuary in Coastal Maine. Rhodora 119: 201-211 (undergraduate student project of Ian Medeiros; PDF)
72. Urban, A. J.**, Mittelhauser, G. H., Dickinson, M.**, and N. Rajakaruna. 2017. The Alpine Vascular Plants of Baxter State Park, Maine, USA. Rhodora 119: 110-131 (PDF; undergraduate senior project of Abbe Urban)
2016
71. Rajakaruna, N. 2016. In Memoriam. Arthur Rice Kruckeberg: Geobotanist Extraordinaire (1920-2016). Madrono 63: 367-370 (PDF)
70. Seneviratne, M., S. Gunaratne, T. Bandara, L. Weerasundara, N. Rajakaruna, H.M.S.P. Madawala, G. Seneviratne, and M. Vithanage. 2016. Plant growth promotion by Bradyrhizobium japonicum under heavy metal stress. South African Journal of Botany 105: 19-24 (PDF)
69. Negoita, L.**, Dickinson, M.**, Mittelhauser, G. H. and Rajakaruna, N. 2016. A Comparative Study of the Flora and Soils of Great Duck and Little Duck Islands, Maine, USA. Rhodora 118: 46-85 (PDF; undergraduate senior project of Luka Negoita)
68. Boyd, R. S., Krell, N. T.*, and Rajakaruna, N. 2016. Extreme Environments. In: Oxford Bibliographies in Ecology. Ed. David Gibson. New York: Oxford University Press; DOI: 10.1093/obo/9780199830060-0152 (PDF)
67. Stern, M.**, Medeiros, I. D.*, Negoita, L.**, and N. Rajakaruna. 2016. Limestone flora of the Simonton Corner Quarry Preserve, Rockport, Maine, USA. Rhodora 118: 206-226 (PDF; undergraduate senior project of Margaret Stern; Supplementary Table)
66. Trau, M.**, Owings, R.**, and N. Rajakaruna. 2016. Implementing Traditional Ecological Knowledge in Conservation Efforts. In Plant Biodiversity: Monitoring, Assessment and Conservation, pp. 525-535, Eds. A. A. Ansari and S. S. Gill. CAB International (PDF)
2015
65. Van der Ent, A., N. Rajakaruna, R. S. Boyd, G. Echevarria, R. Repin, and D. Williams. 2015. Global research on ultramafic (serpentine) ecosystems (8th International Conference on Serpentine Ecology in Sabah, Malaysia). Australian Journal of Botany 63: iii-iv (PDF)
64. Bandara, T., I. Herath, P. Kumarathilaka, M. Seneviratne, G. Seneviratne, N. Rajakaruna, and M.Vithanage. 2015. Role of woody biochar and fungal-bacterial co-inoculation on soil enzyme activity and heavy metal immobilization in serpentine soil. Journal of Soils and Sediments; DOI 10.1007/s11368-015-1243-y (PDF)
63. Van der Ent, A., N. Rajakaruna, R. S. Boyd, G. Echevarria, R. Repin, and D. Williams. 2015. Global Research on the Ecology of Ultramafic (Serpentine) Ecosystems (8th International Conference in Serpentine Ecology, Sabah, Malaysia): a summary and synthesis. Australian Journal of Botany 63: 1-16 (PDF)
62. Burgess, J.L., Szlavecz, K., Rajakaruna, N., and Swan, C.M. 2015. Ecotypic Differentiation of mid-Atlantic Quercus species in response to Ultramafic Soils. Australian Journal of Botany 63: 308-323 (PDF)
61. Seneviratne, M., Seneviratne,G., Madawala, H.M.S.P. Iqbal, M.C.M., Rajakaruna, N. and Vithanage, M. 2015. Role of bacterial-fungal interactions on heavy metal phytotoxicity in serpentine soil. Australian Journal of Botany 63: 261-268 (PDF)
60. Medeiros, I. D.*, Rajakaruna, N. and Alexander, E. B. 2015. Gabbro Soil-Plant Relations in the California Floristic Province. Madrono 62: 75-87 (PDF)
59. Gall, J. E.**, Boyd R. S., and N. Rajakaruna. 2015. Transfer of heavy metals through terrestrial food webs: A review. Environmental Monitoring and Assessment 187: 201. doi:10.1007/s10661-015-4436-3 (PDF)
58. Burgess, J.L., Szlavecz, K., Rajakaruna, N., Lev, S. and Swan, C.M. 2015. Vegetation Dynamics and Mesophication in Response to Conifer Encroachment within an Ultramafic System. Australian Journal of Botany 63: 292-307 (PDF)
57. Ghasemi, R., Z. Zare Chavoshi, R. S. Boyd, and N. Rajakaruna. 2015. Calcium:magnesium ratio affects environmental stress sensitivity in the serpentine endemic Alyssum inflatum Nyár. (Brassicaceae). Australian Journal of Botany 63: 39-46 (PDF)
56. Chathuranga, P. K. D., S. K. A. T. Dharmasena, N. Rajakaruna, and M. C. M. Iqbal. 2015. Growth and nickel uptake by serpentine and non-serpentine populations of Fimbristylis ovata (Cyperaceae) from Sri Lanka. Australian Journal of Botany 63: 128-133 (PDF)
55. Venter, A., A. Levanets, S. Siebert, and N. Rajakaruna. 2015. A preliminary survey of the diversity of soil algae and cyanoprokaryotes on mafic and ultramafic substrates in South Africa. Australian Journal of Botany 63: 341-352 (PDF)
54. Van der Ent, A., N. Rajakaruna, R. S. Boyd, G. Echevarria, and R. Repin. eds. 2015. Ultramafic Ecosytems. Proceedings of the Eighth International Conference on Serpentine Ecology. Australian Journal of Botany 63 (Special Issues 1-4; Parts 1, 2)
2014
53. Rajakaruna, N., R. S. Boyd, and T. B. Harris**. 2014. Synthesis and Future Directions: What have harsh environments taught us about ecology, evolution, conservation and restoration. In Plant Ecology and Evolution in Harsh Environments. Pp. 393-409. Nova Science Publishers, Inc., NY, USA (PDF)
52. Herath, I., P. Kumarathilaka, A. Navaratne, N. Rajakaruna, and M. Vithanage. 2014. Immobilization and phytotoxicity reduction of heavy metals in serpentine soil using biochar. Journal of Soils and Sediments 15: 126-138 (PDF)
51. Medeiros, I. D.*, Fryday, A. M., and N. Rajakaruna. 2014. Additional lichen records and minerological data from metal-contaminated sites in Maine. Rhodora 116: 323-347 (PDF)
50. Rajakaruna, N., R. S. Boyd, and T. B. Harris**, eds. (2014). Plant Ecology and Evolution in Harsh Environments. Nova Science Publishers, Inc., NY, USA. 426p
49. Mansfield, M.*, Pope, N.**, Mittlehauser, G., and N. Rajakaruna. 2014. Diversity and Soil-Tissue Elemental Relations of Vascular Plants of Callahan Mine, Brooksville, Maine, USA. Rhodora 116: 283-322 (PDF; undergraduate senior project of Maggie Mansfield; Supplementary Table)
48. Ghasemi, R., Chavoshi, Z. Z., Boyd, R. S., and N. Rajakaruna. 2014. A preliminary study of the role of nickel in enhancing flowering of the nickel hyperaccumulating plant Alyssum inflatum Nyár. (Brassicaceae). South African Journal of Botany 92: 47-52 (PDF)
47. N. Rajakaruna and R. S. Boyd. 2014. Geoecology. In: Oxford Bibliographies in Ecology. Ed. David Gibson. New York: Oxford University Press (PDF; PDF2)
46. Vithanage, M., Rajapaksha, A. U, Oze, C., Rajakaruna, N. and C.B. Dissanayake. 2014. Metal release from serpentine soils in Sri Lanka. Environmental Monitoring and Assessment, DOI 10.1007/s10661-014-3626-8 (PDF)
45. N. Rajakaruna and R. S. Boyd. 2014. Serpentine Soils. In: Oxford Bibliographies in Ecology. Ed. David Gibson. New York: Oxford University Press (PDF;PDF2)
44. Barton, J.*, Ciccotelli, B.**, Gall, J. E.*, Olday, F. C., Connery, B., T. B. Harris**, A. M. Fryday, and N. Rajakaruna. 2014. Lichens of six vernal pools in Acadia National Park, ME, USA. Evansia 31: 31-39 (PDF; undergraduate senior project of Jason Barton)
43. Neilson, S.** and N. Rajakaruna. 2014. Phytoremediation of Agricultural Soils: Using plants to clean metal-contaminated arable lands. In: Phytoremediation: Management of Environmental Contaminants, pp. 159-168, Eds. A. A. Ansari, S. S. Gill, G. R. Lanza, and L. Newman. Springer (PDF).
2013
42. Boyd, R. S. and N. Rajakaruna. 2013. Heavy Metal Tolerance. In: Oxford Bibliographies in Ecology. Ed. David Gibson. New York: Oxford University Press (PDF; PDF2)
41. Rajakaruna, N. and R. S. Boyd. 2013. Edaphic Factor. In: Reference Module in Earth Systems and Environmental Sciences. Ed. Scott A. Elias. Elsevier, Oxford, United Kingdom (article reproduced from Encyclopedia of Ecology, 2008, Pages 1201–1207; PDF)
40. Gall, J. E.* and N. Rajakaruna. 2013. The physiology, functional genomics, and applied ecology of heavy metal-tolerant Brassicaceae In: Brassicaceae: Characterization, Functional Genomics and Health Benefits (Ed. Minglin Lang), pp. 121-148. Nova Science Publishers, Inc., NY, USA (PDF)
39. Pope, N.**, Fong, M.^, Boyd, R., and N. Rajakaruna. 2013. The role of elevation and soil chemistry in the distribution and ion accumulation of floral morphs of Streptanthus polygaloides Gray (Brassicaceae), a Californian nickel hyperaccumulator. Plant Ecology and Diversity 7: 421-432 (PDF; Supplementary Material; Master’s Thesis research of Mike Fong)
2012
38. Harris, T.B.**, N. Rajakaruna, S.J. Nelson, and P.D. Vaux. 2012. Stressors and Threats to the Flora of Acadia National Park, Maine: Current Knowledge, Information Gaps, and Future Directions. Journal of the Torrey Botanical Society 139: 323-344 (PDF)
37. Rajakaruna, N., Knudsen, K., Fryday, A., O’Dell, R. E., Pope, N.**, Olday, F. C., and S. Woolhouse^. 2012. Investigation of the importance of rock chemistry for saxicolous lichen communities of the New Idria serpentinite mass, San Benito County, California, USA. The Lichenologist 44: 695-714, DOI: http://dx.doi.org/10.1017/S0024282912000205 (PDF)
36. Yost, J. M.^, Barry, T.^, Kay, K.M. and N. Rajakaruna. 2012. Edaphic adaptation maintains the coexistence of two cryptic species on serpentine soil. American Journal of Botany 99 (5): 890-897. (PDF; Master’s Thesis research of Teri Barry)
35. Bieger, A., N. Rajakaruna, and S. P. Harrison. 2012. Little evidence for local adaptation to soils or microclimate in the postfire recruitment of three Californian shrubs. Plant Ecology and Diversity 7: 411-420, DOI:10.1080/17550874.2012.701670 (PDF; Supplementary Materials)
34. Davoodian, N.*, J. Bosworth*, and N. Rajakaruna. 2012. Mycorrhizal colonization of Hypericum perforatum L. (Hypericaceae) on adjacent serpentine and granite outcrops on the Deer Isles, Maine, USA. Northeastern Naturalist 19: 517-526 (PDF; undergraduate senior project of Naveed Davoodian)
33. Neilson, S.** and N. Rajakaruna. 2012. Roles of rhizospheric processes and plant physiology in phytoremediation of contaminated sites using oilseed Brassicas.In: Anjum N. A., Ahmad I., Pereira M. E., Duarte A. C., Umar S., Khan N. A. (Eds.) The Plant Family Brassicaceae: Contribution Towards Phytoremediation, pp. 313-330. Environmental Pollution Book Series, Vol. 21, Springer (Science + Business Media), Dordrecht, The Netherlands (PDF)
2011
32. Anacker, B. L., N. Rajakaruna, D. D. Ackerly, S. P. Harrison, J. E. Keeley, and M. C. Vasey. 2011. Ecological strategies in California chaparral: Interacting effects of soils, climate, and fire on specific leaf area. Plant Ecology and Diversity 4: 179-188, DOI:10.1080/17550874.2011.633573 (PDF; Supplementary Materials).
31. Rajakaruna, N., T. B. Harris*, S. Clayden, A. Dibble, and F. S. Olday. 2011. Lichens of Callahan Mine, a copper and zinc-enriched Superfund site in Brooksville, Maine, U.S.A. Rhodora 113: 1-31 (PDF)
30. Ciccotelli, B.*, T. B. Harris*, B. Connery, and N. Rajakaruna. 2011. A preliminary study of the vegetation of vernal pools of Acadia National Park, Mount Desert Island, Maine, USA. Rhodora 113: 260-279 (PDF; undergraduate senior project of Brett Ciccotelli)
29. R. E. O’Dell and N. Rajakaruna. 2011. Intraspecific variation, adaptation, and evolution. In: Harrison, S. P. and N. Rajakaruna (Eds.). Serpentine: Evolution and Ecology in a Model System, pp. 97-137. University of California Press, Berkeley, CA, USA. (PDF).
28. S. P. Harrison and N. Rajakaruna. 2011. What have we learned from serpentine about evolution, ecology, and other sciences? In: Harrison, S. P. and N. Rajakaruna (Eds.). Serpentine: Evolution and Ecology in a Model System, pp. 417-427. University of California Press, Berkeley, CA, USA. (PDF).
27. Harrison, S. P. and N. Rajakaruna (Eds.). 2011. Serpentine: Evolution and Ecology in a Model System. University of California Press, Berkeley, CA, USA . (See Book Review 1; Book Review 2)
2010
26. Pope, N.*, Harris, T. B.*, and N. Rajakaruna. 2010. Vascular plants of adjacent serpentine and granite outcrops on the Deer Isles, Maine, USA. Rhodora 112:105-141. (PDF; undergraduate senior project of Nate Pope)
2009
25. R. S. Boyd, A. R. Kruckeberg, and N. Rajakaruna. 2009. Biology of ultramafic rocks and soils: research goals for the future. Northeastern Naturalist 16 (5): 422-440. (PDF)
24. Rajakaruna, N. and R. S. Boyd. 2009. Advances in serpentine geoecology: A retrospective. Northeastern Naturalist 16 (5): 1-7. (PDF)
23. Rajakaruna, N. 2009. Serpentinophiles from California and across the world gather in Maine to highlight recent research on soil-biota relations of serpentine outcrops. Fremontia 37 (1): 21-24. (PDF)
22. Rajakaruna, N., and R. S. Boyd (Eds.). 2009. Soil and Biota of Serpentine: A World View. Proceedings of the Sixth International Conference on Serpentine Ecology. Northeastern Naturalist 16 (Special Issue 5).
21. Harris, T. B.* and N. Rajakaruna. 2009. Adiantum viridimontanum, Aspidotis densa, Minuartia marcescens, and Symphyotrichum rhiannon: Additional Serpentine Endemics from Eastern North America. Northeastern Naturalist 16 (5): 111-120. (PDF)
20. Rajakaruna, N., N. Pope*, J. Perez-Orozco*, and T. B. Harris*. 2009. Ornithocoprophilous Plants of Mount Desert Rock, a Remote Bird-Nesting Island in the Gulf of Maine, USA. Rhodora 111: 417-448. (PDF)
19. Rajakaruna, N., T. B. Harris*, and E. B. Alexander. 2009. Serpentine geoecology of eastern North America: a review. Rhodora 111:21-108. (PDF)
18. Briscoe, L. R. E.*, Harris, T. B.* Dannenberg, E.*, Broussard, W.*, Olday, F. C., and N. Rajakaruna. 2009. Bryophytes of adjacent serpentine and granite outcrops on the Deer Isles, Maine, USA. Rhodora 111:1-20. (PDF; undergraduate senior project of Laura Briscoe)
2008
17. Rajakaruna, N. and R.S. Boyd. 2008. The edaphic factor. Pp. 1201–1207, In S.E. Jorgensen and B. Fath (Eds.). The Encyclopedia of Ecology. Volume 2. Elsevier, Oxford, United Kingdom. (PDF)
16. Vaux, P. D., Nelson, S. J., Rajakaruna, N. Mittelhauser, G., Bell, K., Kopp, B., Peckenham, J. and G. Longsworth. 2008. Assessment of natural resources and watershed conditions in and adjacent to Acadia National Park. Natural Resource Technical Report NPS/HTLN/NRTR—2006/001. National Park Service, Fort Collins, Colorado, USA. (PDF)
2007
15. Harris, T. B.*, Olday, F. C., and N. Rajakaruna. 2007. Lichens of Pine Hill, a peridotite outcrop in Eastern North America. Rhodora 109: 430-447. (PDF; undergraduate senior project of Tanner Harris)
2006
14. Bohm, B. A. and N. Rajakaruna. 2006. The Lasthenia californica story: It started with flavonoids. Natural Product Communications 11: 1013-1022. (PDF)
13. Rajakaruna, N., Tompkins, K. M.*, and P. G. Pavicevic.* 2006. Phytoremediation: An affordable green technology for the clean-up of metal contaminated sites in Sri Lanka. Ceylon Journal of Science 35: 25-39. (PDF)
2004
12. Rajakaruna, N. and J. Whitton. 2004. Trends in the evolution of edaphic specialists with an example of parallel evolution in the Lasthenia californica complex. In: Plant adaptation: Molecular Biology and Ecology, Q.C.B. Cronk, J.Whitton , R. Ree and I.E.P. Taylor (Eds), pp. 103-110. (PDF)
11. Rajakaruna, N. and A. J. M. Baker. 2004. Serpentine: A model habitat for botanical research in Sri Lanka. Ceylon Journal of Science 32: 1-19. (PDF)
10. Rajakaruna, N. 2004. The edaphic factor in the origin of species. International Geology Review 46: 471-478. (PDF)
2003
9. Rajakaruna, N. 2003. Edaphic differentiation in Lasthenia: A model for studies in evolutionary ecology. Madrono 50: 34-40. (PDF)
8. Rajakaruna, N., Baldwin, B. G., Chan, R., Desrochers, A. M., Bohm, B. A., and J. Whitton. 2003. Edaphic races and phylogenetic taxa in the Lasthenia californica complex (Asteraceae: Heliantheae): An hypothesis of parallel evolution. Molecular Ecology 12: 1675-1679. (PDF)
7. Rajakaruna, N., Bradfield, G. E., Bohm, B. A., and J. Whitton. 2003. Adaptive differentiation in response to water stress by edaphic races of Lasthenia californica (Asteraceae). International Journal of Plant Sciences 164: 371-76. (PDF)
6. Rajakaruna, N., Siddiqi, M. Y., Whitton, J., Bohm, B. A., and A. D. M. Glass. 2003. Differential responses to Na+/K+ and Ca2+/Mg2+ in two edaphic races of the Lasthenia californica complex (Asteraceae): A case for parallel evolution of physiological traits. New Phytologist 157: 93-103. (PDF)
2002
5. Rajakaruna, N. and B. A. Bohm. 2002. Serpentine and its vegetation: A preliminary study from Sri Lanka. Journal of Applied Botany 76: 20-28. (PDF)
4. Rajakaruna, N., Harris, C. S., and G.H.N. Towers. 2002. Antimicrobial activity of plants collected from serpentine outcrops in Sri Lanka. Pharmaceutical Biology 40: 235-244. (PDF)
2000
3. Whitton, J. and N. Rajakaruna. 2000. Plant biodiversity – an overview. In: S.L. Levin (Ed.). Encyclopedia of Biodiversity, pp. 621-630. Academic Press, San Diego, CA. (PDF; PDF Scanned)
1999
2. Rajakaruna, N. and B.A. Bohm. 1999. The edaphic factor and patterns of variation in Lasthenia californica (Asteraceae). American Journal of Botany 86:1576-1596. (PDF)
1998
1. Bernston, G.M, Rajakaruna, N., and F.A. Bazzaz. 1998. Growth and nitrogen uptake of an experimental community of annuals exposed to elevated atmospheric CO2. Global Change Biology 4: 607-627. (PDF)
Special Journal Issues
Picó, F. X., R. J. Abbott, L. D. Llambi, N. Rajakaruna, A. S. T. Papadopulos, and L. Nagy (eds). 2022. The ecology and evolution of plants in extreme environments. Plant Ecology and Diversity 15 (5-6).
Edited Books
Harrison, S. P. and N. Rajakaruna (Eds.). 2011. Serpentine: Evolution and Ecology in a Model System. University of California Press, Berkeley, CA, USA (See Book Review 1; Book Review 2); Available via Amazon
Rajakaruna, N., R. S. Boyd, and T. B. Harris**, eds. (2014). Plant Ecology and Evolution in Harsh Environments. Nova Science Publishers, Inc., NY, USA. 426p (See Book Review); Available via Amazon
Guest Edited Conference Proceedings
Guillaume Echevarria; Alan J. M. Baker; Robert S. Boyd; Antony van der Ent; Takafumi Mizuno; Nishanta Rajakaruna; Shota Sakaguchi; Aida Bani (Eds.). 2018. Ultramafic Ecosystems: Proceedings of the 9th International Conference on Serpentine Ecology. Ecological Research 33, Issues 3, 4.
Van der Ent, A., N. Rajakaruna, R. S. Boyd, G. Echevarria, and R. Repin. eds. 2015. Ultramafic Ecosytems. Proceedings of the Eighth International Conference on Serpentine Ecology. Australian Journal of Botany 63 (Special Issues 1-4; Parts 1, 2)
Rajakaruna, N., and R. S. Boyd (Eds.). 2009. Soil and Biota of Serpentine: A World View. Proceedings of the Sixth International Conference on Serpentine Ecology. Northeastern Naturalist 16 (Special Issue 5).
Nishanta "Nishi" Rajakaruna 
